Site by Bennett Web & Design Co.
2012 SOUTHEASTERN NATURALIST 11(2):205–218
Effects of Feral Swine (Sus scrofa) on Alligator (Alligator
mississippiensis) Nests in Louisiana
Ruth M. Elsey1,*, Edmond C. Mouton, Jr.2, and Noel Kinler2
Abstract - Rapid spread of the introduced Sus scrofa (Feral Hog) is a major concern for
many landowners and land managers due to its destructive rooting behavior which damages
natural habitats. Feral Swine have also been reported as infrequent predators of
Alligator mississippiensis (American Alligator) eggs, with only seven nests lost in three
prior studies combined (Fogarty 1974, Ruckel and Steele 1984, Woodward et al. 1992).
In response to increasing reports by Louisiana landowners of Alligator nest losses due
to Feral Swine, we sent a questionnaire addressing this issue to licensed Louisiana Alligator
farmers who are permitted to collect eggs from wild nests. Over half (51.4%) of
the farmers reported loss of Alligator nests in 2011; some 590 nests were damaged or
destroyed on 36 separate properties across the state. Four farmers, some of whom have
twenty or more years of experience collecting Alligator eggs, reported this is the first
year in which they have lost nests to Feral Swine. Other farmers reported seeing wild
hogs while in the field or seeing sign of hogs, which suggests future potential losses
may be incurred and that the range and population level of this non-native species is
expanding in important Alligator nesting habitat in Louisiana. Nearly all farmers who
had nests destroyed by Feral Swine (94.7%) reported hog damage is increasing on their
properties. Some farmers reported that hog removal efforts limited their Feral Swine
damage this year relative to past years. In addition to deleterious effects on wetlands
habitats caused by Feral Swine, the financial impact of loss of the Alligator egg revenue
Many landowners across the United States are experiencing problems with
property damage from Sus scrofa L. (Feral Swine), which are expanding their
range (Ditchkoff and Mayer 2009, Mayer and Brisbin 2009). In addition to
agricultural and residential damage, Feral Hogs can destroy crops, damage
wildlife habitat, compete with other species for food, prey on wildlife, and
transmit diseases to wildlife, livestock, and humans (Kimmel 2011, Perot 2011).
They have established populations in at least 38 states and are spreading rapidly,
with estimated economic losses to agriculture and the environment at $800
million annually (Mouton 2009). Additionally, their extensive rooting of soils
can disrupt wetlands and lead to wetlands losses in already fragile ecosystems
(Mouton 2009). Methods are being developed to accurately survey wild pigs
(Williams et al. 2011), and recent legislation has been enacted in Louisiana to
provide more options for property owners to take outlaw quadrupeds, including
1Louisiana Department of Wildlife and Fisheries, 5476 Grand Chenier Highway, Grand
Chenier, LA 70643. 2Louisiana Department of Wildlife and Fisheries, 2415 Darnall Road,
New Iberia, LA 70560. *Corresponding author - firstname.lastname@example.org.
206 Southeastern Naturalist Vol. 11, No. 2
Feral Hogs (Louisiana Department of Wildlife and Fisheries press release, 14
July 14 2011; Perot 2011). Studies are underway in Louisiana to evaluate the
possible effect of soil quality on reproductive efficiency in this prolific species
(O’Boyle and Tolson 2011) that may help elucidate why their range is expanding
in certain regions, and a recent report (Mayer and Brisbin 2009) reviews
management and control techniques for this damaging species. Although Feral
Hogs have only rarely been noted as a predator of Alligator mississippiensis
Daudin (American Alligator; hereafter just Alligator) eggs in Louisiana (McIlhenny
1935), their recent population growth and range expansion could make
this mortality factor of greater import adversely affecting Louisiana’s valuable
Numerous studies have been conducted on the nesting ecology of Alligators
in southeastern states, and wildlife managers in Louisiana, Florida, South
Carolina, Texas, and Georgia have documented nest losses due to flooding and
predation (Cooper and Slaughter 2008, Deitz and Hines 1980, Fleming et al.
1976, Goodwin and Marion 1978, Hunt and Ogden 1991, Joanen 1969, Metzen
1977, Platt et al. 1995, Ruckel and Steele 1984, Wilkinson 1983). Procyon lotor
L. (Raccoons) are the most often cited mammalian predator in these studies, but
in some cases Lontra canadensis Schreber (River Otters) have been suspected
as predators on Alligator eggs (Deitz and Hines 1980), and Ursus americanus
Pallas (American Black Bears) have also been documented as causing nest/egg
loss (Hunt and Ogden 1991, Metzen 1977).
In McIlhenny’s (1935) classic study on Alligators, he noted Alligator eggs to
be eaten by a number of animals, including “’coons, opossums, skunks, hogs,
and bears”. Fogarty (1974) noted a hog was seen rooting through one of 64
nests (1.6%) studied in the Florida Everglades. More recently, Ruckel and Steele
(1984) noted that four of 31 nests at their Rhetts Island study site in Georgia were
destroyed by Feral Hogs, and one other nest was destroyed by a combination of
Raccoons and Feral Hogs; thus, 5 of 31 nests (16.1%) were lost all or in part to
Feral Hogs. In a long-term, multi-year study in Florida, Woodward et al. (1992)
noted one nest was depredated by Feral Hogs, as evidenced by extensive rooting
and hoof-prints around the nest. Biological staff from our agency noted signs
of hog presence in nesting areas and nests destroyed by hogs in 1997 during a
pilot study evaluating Alligator nests in swamp habitat with a cooperating eggcollection
permittee (Campbell 1997).
Commercial Alligator farming programs exist in many southeastern states, including
Louisiana. A major component of the program in Louisiana involves egg
“ranching”, wherein Alligator farmers may collect eggs from nests in the wild to
stock Alligator farms under permits and quotas established by the Louisiana Department
of Wildlife and Fisheries (LDWF). This program has been described in
detail (Elsey et al. 2001), and can have an economic value of nearly $60 million
in strong market years for the valuable leather industry (LDWF 2009).
To better evaluate Alligator nest losses and quantify potential impact to Alligator
nesting habitat in Louisiana due to Feral Hogs, we developed a brief
2012 Ruth M. Elsey, Edmond C. Mouton, Jr., and Noel Kinler 207
questionnaire to request information on nest destruction due to this species. Information
obtained might be of use in assisting wildlife managers in controlling
Feral Hogs on their properties which may damage valuable wetland resources,
including Alligator eggs.
A brief (one page) questionnaire was developed and sent to licensed Alligatoregg
permittees (licensed Louisiana Alligator farmers, who have permits to collect
Alligator eggs from the wild). Thirty-seven farmers have such egg permits and
were mailed the questionnaire. Self-addressed stamped return envelopes were provided
to facilitate replies to the brief survey, and a cover letter assured respondents
their replies would in no way affect future egg quotas which are determined by our
agency. One Alligator farmer/permittee was coincidentally in the LDWF office on
the day the questionnaire was to be mailed; he was provided his copy in person. Another
Alligator farmer had provided some information on Feral Hog damage when
an informal inquiry was sent to him via e-mail; he was not mailed another form, and
details were obtained in follow up e-mail correspondence and by telephone.
The form also indicated farmers could telephone the LDWF office and discuss
their answers and observations over the telephone, rather than having to complete
the written questionnaire, if this was easier. The brief questions included:
1. Did you see any evidence of hog (Feral Pig) damage to Alligator nests
during your collections this year?
2. If so, on which land companies did you see hog damage to Alligator
nests? (please list how many nests damaged on each property; and whether
they were slightly damaged or completely destroyed). Please provide as
much detail as is possible. Any photos are appreciated as well.
3. Have you ever seen hog damage to Alligator nests in past years? List
years if possible.
4. Does hog damage seem to be increasing on your properties, decreasing,
or staying about the same?
5. Have you observed live hogs in the marsh when collecting eggs or flying
to mark nests? If so, which properties?
6. Have you seen hog sign (footprints/tracks, scats/droppings, etc.) near
nests while collecting nests?
7. How many years have you been collecting eggs from the wild?
The questionnaire was mailed in mid-July (July 19), by which time a few
farmers may have already recently completed the summer’s egg collections,
but many would still be actively collecting eggs. Thus, we hoped most farmers
would remember if they had seen any evidence of Feral Swine damage, but we
did not want to seem to be “encouraging” reporting of questionable findings or
cause over interpretation of damage to Alligator nests caused by other means.
The number of licensed egg permittees is low (n = 37), and we had a 100%
response rate. In some cases, this was done via telephone and we recorded
the information supplied.
208 Southeastern Naturalist Vol. 11, No. 2
We were confident Alligator farmers could distinguish nest depredation
by Feral Swine from other predators (mainly Raccoons), as nests depredated
by Raccoons typically have numerous small penetration holes, apparently made
Figure 1. An Alligator nest depredated by Raccoons. The nest damaged by Raccoons is
mostly structurally intact, with eggshells scattered about the exterior of the nest.
Figure 2. A typical undamaged Alligator nest. The typical nest has eggs concealed within
the inner egg cavity.
2012 Ruth M. Elsey, Edmond C. Mouton, Jr., and Noel Kinler 209
as Raccoons probed for eggs (Figs. 1, 2; Woodward et al. 1992). Feral Hogs
would generally create more nest damage and disrupt the integrity of the nest
mound with rooting activities, and hoof prints and tracks of pigs might be seen
at the nest site (Fig. 3). Additionally, nearly 68% of licensed Louisiana Alligator
farmers have over twenty years of experience collecting Alligator eggs and thus
were felt to be reliable observers in submitting the requested data.
Figure 3. Photo showing an Alligator nest damaged by Feral Swine. Note the extensive
damage and disruption of nest integrity caused by Feral Swine. Photograph © Jeff Donald.
210 Southeastern Naturalist Vol. 11, No. 2
The survey was not developed until after the egg collection season started;
thus, the results in this study are in many cases estimates the farmers made after
completing egg collections. If a farmer reported a range of nests damaged (for
example, “3–4 nests lost on Property A”), we would assume the lesser quantity
and consider three nests lost. If they reported a range such as “8–10 nests lost on
Property B”, we considered this as nine nests lost. The farmer with the most nests
damaged reported an estimate based on a percentage of nests damaged in each
section that the ultra-light aircraft pilot surveyed, and the number of nests ultimately
marked (the pilot did not throw a nest marker at nests already destroyed
by Feral Swine at the time of survey, as ground crews would not need to visit that
individual nest site).
Some farmers specifically reported they did not have hog damage but did lose
a few nests to Raccoons; others reported some Feral Swine damage but more Raccoon
damage, which lent support to our supposition that Alligator farmers could
distinguish the two factors. One farmer reported rare Alligator nest losses due to
bears; the property manager has frequently observed bears on the area but has not
Hundreds of Alligator egg permits are issued to licensed Louisiana Alligator
farmers annually; in 2011, 531 permits were issued to individual properties with a
total egg quota total of over 785,000 eggs. The number of egg permits individual
farmers have varies widely (range = 1–75); the number of eggs in each farmer’s
quota total ranges from 700 eggs to 95,915 eggs. The single smallest permit
has an egg quota of 35 eggs, and the largest single permit has a quota of 55,535
eggs. Alligators occur statewide in Louisiana, but the majority of the Alligator
habitat (and thus Alligator nests, population, and egg quotas) occur in the coastal
parishes. A variety of habitats exist, including large lakes, upland areas, cypress
tupelo swamps, and coastal marshes (Elsey and Kinler 2004).
Alligator nest damage in 2011, extent of nest damage, and properties affected
Numerous farmers (51.4%) reported Feral Swine damage to Alligator nests
in 2011. Nineteen farmers reported losing approximately 590 Alligator nests to
Feral Swine in 2011. Properties affected were broadly distributed across the state
(Fig. 4), although more occurred in the coastal zone where Alligator populations
are greatest. Two farmers who collect eggs on one large property lost some 315
nests as a conservative estimate for their combined collections. They successfully
collected 40,500 eggs combined; but lost approximately 25% of the nests
produced to Feral Hogs. One farmer with 75 egg permits lost 132 nests on nine
properties; fifty nests were lost on his largest property, which had about one third
of his entire egg quota. One farmer lost 27 nests on a single large wetland property;
another farmer with a single egg permit lost 10 of 39 (25.6%) Alligator nests
he located to hog damage. In 2011, Louisiana Alligator farmers collected over
352,000 eggs from some 13,000 nests. Thus, the estimated 600 nests damaged by
hogs represents approximately 4–5% of the nests collected.
2012 Ruth M. Elsey, Edmond C. Mouton, Jr., and Noel Kinler 211
Some farmers reported damage on only one property; others noted hog damage
to multiple properties. Nest quantities reported damaged ranged from one to
approximately 300. Most farmers reported damaged nests were totally destroyed;
one farmer verbally reported the nests were “flattened”. Only one farmer noted
that in some cases they were able to collect some eggs from nests partially damaged
by Feral Swine. Another farmer reported the top of the nest was torn away
and a few eggs consumed; the other eggs were scattered and lost due to heat exposure.
The damaged or destroyed nests were located on 36 separate properties
across the state (Fig. 4).
One farmer noted hog damage to nests on seven different properties in 2011
(approximately 15 nests lost). His most severely affected property might have
been damaged even more extensively had it not been for the trapping and removal
of 69 Feral Swine this year. Another farmer removed over 50 pigs by trapping;
he has seen hogs and hog sign near nests for many years, but this is the first year
he actually lost two nests to rooting by hogs.
Figure 4. Map showing parish locations with hog damage in Louisiana in 2011. Each
point represents a property with at least one nest damaged by Feral Swine and indicates
the parish wherein that property is located. Points located on the border of two parishes
represent land properties with hog damage with sufficient acreage that the properties are
located within two adjoining parishes.
212 Southeastern Naturalist Vol. 11, No. 2
Prior Alligator nest damage and trends in Alligator nest damage
Most farmers who saw damage in 2011 had seen damage previously over the
last few years. One farmer in southwest Louisiana had a decreased nesting effort
in 2011 due to severe drought; thus, he did not attempt egg collections and could
not asses hog damage, but he had previously seen hog damage to Alligator nests
on several properties. Another farmer similarly affected by drought saw so few
nests he did not collect eggs in 2011 in southwest Louisiana, but he previously
had seen Feral Swine damage in southeast Louisiana. Four farmers noted nest
damage attributed to Feral Swine in 2011, but had not seen this previously in the
15 to 22 years experience each had collecting Alligator eggs, indicating Feral
Hogs may be just now extending their ranges in coastal Louisiana and/or increasing
the overall population level.
Nearly all farmers (94.7%) who reported Feral Swine damage in 2011 noted
that hog damage was increasing based on their observations. Several farmers
noted they would previously observe rare hog damage in isolated locations affecting
one or two nests, but now see “areas” with larger regional damage of
several nests. Another farmer similarly described “hot spots” of extensive hog
damage, and said Feral Swine damage seems to be worse in each successive year
over the last three or four years.
One farmer noted that they were seeing less hog damage due to their eradication
program, but the overall range of hogs on their property had increased. This
farmer/land manager had extensive records, and noted a yearly increase in hog
damage, beginning with approximately ten nests lost to Feral Swine in 1998 to
approximately 100 nests lost in 2008. Due to their hog eradication program, they
lost fewer nests (n = 27) in 2011 to hog damage. Thus, three farmers (two others
as noted above in “extent of nest damage”) described beneficial effects of hog
eradication programs in limiting swine damage to Alligator eggs/nests.
Three farmers who did not lose Alligator nests to Feral Swine in 2011 did
report that they saw an increase in hog damage to their properties. One farmer
verbally reported a “population explosion” of Feral Swine causing problems
to him as well as local corn farmers. One farmer said hog damage was decreasing,
but this is based on having had one nest destroyed three years ago
and none since.
Observations of live Feral Swine and hog sign
Most farmers who saw egg damage to nests reported having seen live hogs
both while collecting eggs or when flying to mark nests for later collection by
airboat. One farmer spontaneously remarked he saw Feral Swine “by the dozens”
while conducting egg-collecting activities. Several farmers noted their pilots (helicopter
or ultra-light aircraft) observed live hogs or nest damage while surveying
to locate nest sites; some saw nests already destroyed and thus did not mark those
nests for later collection. Five farmers who had not yet experienced nest loss/
damage to hogs (or had no damage this year, but did in the past) saw live hogs on
their properties while collecting eggs or marking nests.
2012 Ruth M. Elsey, Edmond C. Mouton, Jr., and Noel Kinler 213
As expected, farmers who incurred nest losses to Feral Swine reported seeing
hog sign at nests. Of additional concern is that four farmers reported seeing
hog sign (tracks, rooting) on their properties (but no damage to nests as of yet),
indicating they may have the potential to suffer nest losses as well as continued
wetlands disruption or erosion in future years.
Experience level of respondents
Most farmers who are currently active have been licensed Alligator farmers for
many years; many since the inception of the Alligator egg “ranching” program in
1986. Twenty-three of 34 (67.65%) farmers who responded to this question had
over twenty years of experience and observations. Only four respondents had less
than ten years experience collecting eggs, with the least experienced surveyed
farmer having four years of egg-collecting experience. Thus, in total, the farmers
surveyed have an extensive experience level and have observed a past lack of
Feral Swine and/or seen the influx of this species over the years.
The very high numbers of total nests now being lost and large proportion of
Alligator farmers losing valuable Alligator nests and eggs to Feral Hogs was an
unanticipated result and cause for some concern. A disturbing trend was also
noted of most farmers reporting increasing evidence of hog presence and damage
to their wetlands, Alligator habitat, and egg resources. Several experienced Alligator
farmers saw hog damage for the first time in 2011, suggesting populations
of this non-native species are rising and/or expanding their range in Louisiana.
The results presented here are minimums. Some pilots did not mark nests that
they saw from the air were already destroyed; thus, ground crews would not have
encountered these nests and they would have been unreported as having been
damaged by hogs. Also, some remaining uncollected nests (if any were left after a
farmer collected his full egg quota on a property) may have been later depredated
by Feral Swine.
Early egg collection of Alligator nests can prevent flooding, predation, and
lightning fire losses; avoiding high natural mortality was a factor considered in
implementing egg-collecting programs in Louisiana (Elsey and Trosclair 2008).
Avoidance of Feral Hog damage could also be a reason to encourage participation
in egg-collecting programs in some regions. One farmer suggested his early egg
collections (late June) averted hog losses seen the prior year when he had collected
later than usual (mid-July). Farmers who have “hot spots” with extensive
hog presence should perhaps be encouraged to collect these areas first to avoid
nest losses. It may well be that Feral Swine damage to wild Alligator nests may
have been far greater if the egg-collecting program were not in place, which limits
some of the natural mortality factors.
It may also be that unusually high water levels in 2011 (Mississippi River
levels were much higher than normal due to record snowfall in northern states)
led to increased hog damage this year. In contrast, however, Fleming et al. (1976)
214 Southeastern Naturalist Vol. 11, No. 2
reported an absence of predation by Raccoons on Alligator nests after prolonged
high marsh water levels. Mazzotti (1989) attributed low levels of Crocodylus
acutus Cuvier (American Crocodile) nest predation to the low density of active
nests in any given year. It may be that the higher than average Alligator nesting
in southeast Louisiana in 2011 led to the observed increase in Feral Swine damage
seen this year, in addition to what clearly appears to be range and population
expansion of Feral Swine statewide.
Six farmers noted damage on other properties in past years that were not
damaged this year; it would be of interest to determine why some properties are
targeted or if Feral Swine move vast distances regularly and thus affect Alligator
nests randomly in various locations.
Nest losses have also occurred in other crocodilian species due to pigs; Webb
et al. (1983) noted Feral Hogs took five nests at one site (6% of the total) in a
study on Crocodylus johnstoni Krefft (Australian Freshwater Crocodile) nesting.
Similarly, Hall and Johnson (1987) documented losses of nests of Crocodylus
novaeguineae Schmidt (New Guinea Crocodile) due to predation by Varanus
sp. Merrem (varanid lizards) and Feral Hogs. Platt et al. (2008) noted a nest of
Crocodylus moreletii Dumeril, Bibron, and Dumeril (Morelet’s Crocodile) was
destroyed by Tayassu tajacu L. (Collared Peccary) in northern Belize. Crawshaw
and Schaller (1979) suggested possible local vertebrate predators in their study
of Caiman crocodilus yacare Daudin (Yacare Caiman) in Brazil were Sus scrofa
domesticus L. (Domestic Pigs) and Tayassu (peccaries), among others. Campos
(1993) also documented tracks and feces of Feral Hogs near nests of Yacare
Caiman that had been destroyed by predators in the Pantanal of Brazil. Larriera
and Pina (2000) also found Feral Hog destruction of Caiman latirostris Daudin
(Broad-nosed Caiman) nests in northern Argentina.
The apparent recent increase in Alligator nest losses due to Feral Swine is a disturbing
trend that should be continued to monitored closely, but we do not believe
this is an imminent threat to Alligator populations. However, this additional mortality
factor could become problematic if it continues to increase unchecked; and
we anticipate continued collection of hog-damage data in future years.
In recent years, our aerial nesting surveys have estimated over 30,000 nests
just in coastal Louisiana; thus, the nearly 600 nests lost to Feral Swine would
represent some 2% of the statewide nest production. The possible financial losses
due to Alligator egg destruction by pigs to Alligator industry personnel could be
significant. If eggs are valued at $10 each and using conservative estimates of
25 eggs per nest, a loss of $147,500 would be incurred for just the egg value;
there would be an additional loss of revenue from the finished leather-goods
products. Although this study is limited in scope to Alligator nest damage, of
greater concern is the negative impact these invasive Feral Swine are having by
damaging wetlands along the coast of Louisiana that are recovering from damage
by Myocastor coypus Molina (Nutria) (Mouton 2009). Indeed, researchers
have determined that Feral Swine damage to coastal wetlands is more severe than
Nutria damage (Mouton 2009).
2012 Ruth M. Elsey, Edmond C. Mouton, Jr., and Noel Kinler 215
Many mammalian predators prey upon Louisiana’s resident Anas fulvigula
Ridgway (Mottled Duck), affecting nesting females, ducklings, adults during remigial
molt, and eggs (Moorman and Gray 1994). Feral Hogs have the potential
to prey upon this species, which is of concern due to uncertain population status,
and hunter bag limits have recently been decreased from three Mottled Ducks per
day to one in Louisiana. Feral Hogs have also been reported to destroy Meleagris
gallopavo L. (Wild Turkey) nests (Perot 2011), another popular game species in
Louisiana. Other concerns noted by two Alligator farmers were Feral Swine damage
to local corn farmers, one egg collector reported sugar cane farmers are also
seeing adverse effects due to Feral Swine damage.
As far back as 1929 (Kellogg 1929), the range of swine and Alligators have
apparently overlapped in Louisiana, as a hog was reported found in an Alligator
stomach on Rainey Wildlife Refuge in Vermilion Parish. McIlhenny (1935)
also describes witnessing Alligators taking hogs and details one instance of a
“duroc boar hog” weighing not less than 500 pounds being taken by a large Alligator
while the hog was swimming across a stream. O’Neil (1949) noted hogs
as a predator of Ondatra zibethicus L. (Muskrats) in Louisiana coastal marshes.
However, Alligators preying on hogs is relatively uncommon; Neill (1971)
said “troops of Feral Hogs, so common in parts of the Southeast, are not often
menaced by Alligators.” Taylor (1986) noted hogs in two of 111 Alligator stomachs
(1.8%) examined. In Georgia, Shoop and Ruckdeschel (1990) noted Feral
Swine in five of 28 Alligator fecal samples from scattered localities on Cumberland
Although the finding of Alligator nest losses is not new, and the range of
Alligators and swine in Louisiana is known to overlap, the recent tremendous
increase in extent of damage, apparent Feral Hog range expansion, and potential
for future continued losses are of concern. Land managers and industry
participants may work together with regulatory agencies to consider eradication
programs in the future. Those farmers and land managers who saw live
hogs or signs of hog damage to wetlands (despite them not yet having experienced
actual Alligator nest losses) might be encouraged to consider eradication
in the future to avoid later costly loss of Alligator nests and to collect nests in
those areas as early as possible to avoid Feral Swine damage. The role of the
female Alligator in nest defense as a deterrent to hog depredation could be an
area for future investigation.
The Feral Swine may be another example of a non-native introduced species
which has led to serious unanticipated consequences. It was encouraging to
receive reports that several proactive farmers or land managers who trapped or
removed hogs believed it did limit even further Alligator nest damage than was
reported. Future research might help elucidate if there is a habitat type preferred
by hogs where the most nest damage occurs or if they are generalists and adapt
to many habitat types. There may be landscape characteristics that lead to more
depredation upon Alligator nests by hogs in some regions.
216 Southeastern Naturalist Vol. 11, No. 2
We thank all Alligator farmers/ranchers and land managers who supplied information
and responded to the questionnaire. We thank Drs. David R. Chalcraft and I. Lehr Brisbin,
Jr. for helpful comments to improve the manuscript.
Campbell, L. 1997. In-house memorandum. Photographs from Alligator research. 30
Campos, Z. 1993. Effect of habitat on survival of eggs and sex ratio of hatchlings of Caiman
crocodilus yacare in the Pantanal, Brazil. Journal of Herpetology 27(2):127–132.
Cooper, A., and M.J. Slaughter. 2008. (Abstract). Nesting success of American Alligators
in a southeast Texas coastal marsh. Proceedings of the Annual Conference of Southeastern
Association of Fish and Wildlife Agencies 62:204.
Crawshaw, P., and G.B. Schaller. 1979. Nesting of Paraguyan Caiman (Caiman yacare)
in Brazil. Report No. 11 to Instituto Brasileiro de Desenvolvimento Florestal (IBDF)
and Fundação Brasileira para a Conservação de Natureza, Brasilia, Brazil. 21 pp.
Deitz, D.C., and T.C. Hines. 1980. Alligator nesting in North-central Florida. Copeia.
Ditchkoff, S.S., and J.J. Mayer. 2009. Biology of Wild Pigs: Wild Pig food habits. Pp.
105–143, In J.J. Mayer and I.L. Brisbin, Jr. (Eds.). Wild Pigs: Biology, Damage, Control
Techniques, and Management. Publication SRNL–2009-00869 of the Savannah
River National Laboratory, Aiken, SC.
Elsey, R.M., and N. Kinler. 2004. Louisiana’s Alligator program: Adapting management
as populations recover and risk of unsustainable use decreases. Pp. 92–101, In Crocodiles.
Proceedings of the 17th Working Meeting of the Crocodile Specialist Group,
IUCN - The World Conservation Union, Gland, Switzerland, and Cambridge, UK.
Elsey, R.M., and P.L. Trosclair III. 2008. Effect of timing of egg collection on growth in
hatchling and juvenile American Alligators. Herpetological Bulletin 105:13–18.
Elsey, R.M., L. McNease, and T. Joanen. 2001. Louisiana’s Alligator ranching program:
A review and analysis of releases of captive-raised juveniles. Pp. 426–41, In G. Grigg,
F. Seebacher, and C.E. Franklin (Eds.). Crocodilian Biology and Evolution. Surrey
Beatty and Sons, Chipping Norton, New South Wales, Australia.
Fleming, D.G., A.W. Palmisano, and T. Joanen. 1976. Food habits of coastal marsh
Raccoons with observations of Alligator nest predation. Proceedings of the Annual
Conference of Southeastern Association of Fish and Wildlife Commissioners
Fogarty, M.J. 1974. The ecology of the Everglades Alligator. Pp. 367–374, In P.J. Gleason
(Ed.). Environments of South Florida: Present and Past. Memoir 2: Miami Geological
Society, Miami, FL. 452 pp.
Goodwin, T.H., and W.R. Marion. 1978. Aspects of the nesting ecology of American Alligators
(Alligator mississippiensis) in north-central Florida. Herpetologica 34:43–47.
Hall, P.M., and D.R. Johnson. 1987. Nesting ecology of Crocodylus novaeguineae in
Lake Murray District, Papua New Guinea. Herpetologica 43(2):249–258.
Hunt, R.H., and J.J. Ogden 1991. Selected aspects of the nesting ecology of American
Alligators in the Okefenokee Swamp. Journal of Herpetology 25(4):448–453.
Joanen, T. 1969. Nesting ecology of the Alligator in Louisiana. Proceedings of the Annual
Conference of Southeastern Association of Fish and Wildlife Commissioners
2012 Ruth M. Elsey, Edmond C. Mouton, Jr., and Noel Kinler 217
Kellogg, R. 1929. The habits and economic importance of Alligators. US Department of
Agriculture, Washington, DC. Technical Bulletin No. 147. 36 pp.
Kimmel, F. 2011. Hogs: Problem porkers. Louisiana Wildlife Insider. Spring-summer
2011. Pp. 15, 19.
Larriera, A., and C. Pina. 2000. Caiman latirostris (Broad-Snouted Caiman) nest predation:
Does low rainfall facilitate predator access? Herpetological Natural History
Louisiana Department of Wildlife and Fisheries (LDWF). 2009. 2008–2009 annual report.
Baton Rouge, LA. 98 pp.
LDWF. 2011. Legislation provides more options for property owners contending with
outlaw quadrupeds, nuisance animals. LDWF press release, 14 July 2011. Available
online at http://www.wlf.louisiana.gov. Accessed on 14 July 2011.
Mayer, J.J., and I.L. Brisbin, Jr. 2009. Wild Pigs: Biology, Damage, Control Techniques,
and Management. Publication SRNL–2009-00869 of the Savannah River National
Laboratory, Aiken, SC.
Mazzotti, F. 1989. Factors affecting the nesting success of the American Crocodile, Crocodylus
acutus, in Florida Bay. Bulletin of Marine Science 44(1):220–228.
Metzen, W.D. 1977. Nesting ecology of Alligators on the Okefenokee National Wildlife
Refuge. Proceedings of the Annual Conference of Southeastern Association of Fish
and Wildlife Agencies 31:29–32.
McIlhenny, E.A. 1935. The Alligator’s Life History. The Christopher Publishing House,
Boston, MA.117 pp.
Moorman, T.E., and P.N. Gray. 1994. Mottled Duck (Anas fulvigula). Pp, 1–20, In A.
Poole and F. Gill (Eds.) The Birds of North America. No. 81. The Academy of Natural
Sciences, Philadelphia, PA; The American Ornithologist’s Union, Washington, DC.
Mouton, E. 2009. Testimony before the United States Senate, Committee on Environment
and Public Works, Subcommittee on Water and Wildlife. 3 December 2009. 9 pp.
Neill, W.T. 1971. The food of the Alligator. Pp. 233–249, In The Last of the Ruling Reptiles.
Alligators, Crocodiles, and their Kin. Columbia University Press, New York,
NY. 486 pp.
O’Boyle, M.L., and K.M. Tolson. 2011. The biological potential of the Feral Pig (Sus
scrofa) in Louisiana. (Abstract). Louisiana Association of Professional Biologists
2011 Fall Symposium, Lafayette, LA.
O’Neil, T. 1949. The Muskrat in the Louisiana Coastal Marshes. Louisiana Department
of Wild Life and Fisheries. Federal Aid Section–Fish and Game Division, New Orleans,
LA. 152 pp.
Perot, M. 2011. Coping with Feral Hogs. Landowners for Wildlife, Private Lands Program.
Publication of the Louisiana Department of Wildlife and Fisheries. New Orleans,
LA. 4 pp.
Platt, S.G., R.W. Hastings, and C.G. Brantley. 1995. Nesting ecology of the American
Alligator in Southeastern Louisiana. Proceedings of the Annual Conference of Southeastern
Association of Fish and Wildlife Agencies 49:629–639.
Platt, S.G., T.R. Rainwater, J. B. Thorbjarnarson, and S.T. McMurry. 2008. Reproductive
dynamics of a tropical freshwater crocodilian: Morelet’s Crocodile in northern Belize.
Journal of Zoology 275:177–189.
Ruckel, S.W., and G.W. Steele. 1984. Alligator nesting ecology in two habitats in Southern
Georgia. Proceedings of the Annual Conference of Southeastern Association of
Fish and Wildlife Agencies 38:212–221.
218 Southeastern Naturalist Vol. 11, No. 2
Shoop, C.R., and C.A. Ruckdeschel. 1990. Alligators as predators on terrestrial mammals.
American Midland Naturalist. 124:407–412.
Taylor, D. 1986. Fall food habits of adult Alligators from cypress lake habitat, Louisiana.
Proceedings of the Annual Conference of Southeastern Association of Fish and
Wildlife Agencies. 40:338–341.
Webb, G.J.W., R. Buckworth, and S.C. Manolis. 1983. Crocodylus johnstoni in
the McKinaly River, N.T, VI.* Nesting Biology. Australian Wildlife Research.
Wilkinson, P.W. 1983. Nesting ecology of the American Alligator in coastal South
Carolina. Study Completion Report. August 1978–September 1983. South Carolina
Wildlife and Marine Resources Department, Division of Wildlife and Freshwater
Fisheries, Charleston, SC. 114 pp.
Williams, B.L., R.W. Holtfreter, S.D. Ditchkoff, and J.B. Grand. 2011. Efficiency of
time-lapse intervals and simple baits for camera surveys of Wild Pigs. Journal of
Wildlife Management 75(3):655–659.
Woodward, A.R., C.T. Moore, and M.F. Delany. 1992. Experimental Alligator harvest.
Final Report. Study Number: 7567. Study Period: 1 July 1981–30 June 1991. Bureau
of Wildlife Research. Florida Game and Fresh Water Fish Commission, Gainesville,
FL. 118 pp.